David A. Orwig

Harvard Forest
Harvard University

Petersham, MA 01366
978-724-3302 ext. 250
978-724-3595 (fax)
orwig@fas.harvard.edu

Education

Rochester Institute of Technology	Biology	B.S.	1989
The Pennsylvania State University	Ecology	M.S.	1991
The Pennsylvania State University	Forest Resources	Ph.D.	1995
Harvard University, Harvard Forest	Postdoctoral Research Associate		1995-97

Employment

June 1997 - present. Forest Ecologist. Harvard University, Harvard Forest.
June 1995- June 1997. Postdoctoral Research Associate. Harvard University, Harvard Forest.
Aug. 1989 to May 1995. Graduate Teaching and Research Assistant. The Pennsylvania State University.

Professional Organizations

Ecological Society of America
Torrey Botanical Club
The Tree-ring Society
Hemlock Woolly Adelgid (HWA) Working Group

Research Interests

My research interests encompass all aspects of forest ecology, with particular emphasis on old-growth forest dynamics, dendroecology, and the role of land use history and disturbance on forest structure and composition. These approaches have recently been used to examine forest damage, environmental changes, and related ecosystem processes associated with hemlock woolly adelgid outbreaks in New England hemlock forests.

Refereed Publications

Seischab, F.K. and D. Orwig. 1991. Catastrophic disturbances in the presettlement forests of western New York. Bull. Torrey Bot. Club 118: 117-122.

Orwig, D.A. and M.D. Abrams. 1991. Woody vegetation and fuel survey for evaluating wildfire hazard in Spotsylvania Battlefield. Tech. Rep. NPS/MAR/NRTR-91/052, Philadelphia, PA. 108 pp.

Orwig, D.A. and M.D. Abrams. 1993. Temperate forests of the eastern United States. In S.K. Majumdar, E.W. Miller, D.E. Baker, E.K Brown, J.R. Pratt, and R.F. Schmalz, eds., Conservation and Resource Management. Pennsylvania Academy of Science. pp. 97-116.

Orwig, D.A. and M.D. Abrams. 1994. Land-use history (1720 - 1992), composition, and dynamics of oak-pine forests within the Piedmont and Coastal Plain of northern Virginia. Can. J. For. Res. 24: 1216-1225.

Mikan, C.J., D.A. Orwig, and M.D. Abrams. 1994. Composition, structure and successional dynamics of a pre-European settlement dry oak forest in southeastern Pennsylvania. Bull. Torrey Bot. Club 121: 13-23.

Orwig, D.A. and M.D. Abrams. 1994. Contrasting radial growth and canopy recruitment patterns in Liriodendron tulipifera and Nyssa sylvatica: gap obligate versus gap-facultative tree species. Can. J. For. Res. 24: 2141-2149.

Abrams, M.D. and D.A. Orwig. 1994. Temperate Hardwoods. In Encyclopedia of Agricultural Sciences. Academic Press, Inc. pp. 289-302.

Abrams, M.D., D.A. Orwig, and T. DeMeo. 1995. Dendroecology and successional dynamics of a presettlement-origin white pine-mixed oak forest in the southern Appalachians. Journal of Ecology. 83:123-133.

Abrams, M.D. and D.A. Orwig. 1995. Community and radial growth dynamics of a 320 year-old pitch pine rock outcrop community in southeastern New York. Oecologia. 101:353-360.

Orwig, D.A. and M.D. Abrams 1995. Dendroecological and ecophysiological analysis of gap environments in mixed-oak understories of northern Virginia. Functional Ecology 9:799-806.

Abrams, M.D. and D.A. Orwig. 1996. A review of dendroecology and successional dynamics of old-growth and second-growth oak and pine forests in the eastern United States. Radiocarbon In Tree rings, environment, and humanity (eds. J.S. Dean, D.M. Meko, and T.W. Swetnam) RADIOCARBON 1996, pp. 343-352.

Abrams, M.D. and D.A. Orwig. 1996. A 300-year history of disturbance and canopy recruitment for co-occurring white pine and hemlock on the Allegheny Plateau, U.S.A. Journal of Ecology 84:353-363.

Foster, D.R., D.A. Orwig, and J.S. McLachlan. 1996. Ecological and conservation insights from reconstructive studies of temperate old-growth forests. Trends in Ecology and Evolution 11:419-424.

Abrams, M.D., D.A. Orwig, and M.J. Dockry. 1997. Dendroecology and successional status of two contrasting old-growth oak forests in the Blue Ridge Mountains, U.S.A.. Can. J. For. Res. 27:994-1002.

Orwig, D.A. and M.D. Abrams. 1997. Variation of radial growth responses to drought among species, site, and canopy strata. Trees 11:474-484.

Orwig, D.A. and D.R. Foster. 1998. Forest response to the introduced hemlock woolly adelgid in southern New England, USA. J. Torrey Bot. Soc. 125:59-72.

Orwig, D.A. and M.D. Abrams. 1999. Impacts of early selective logging on the dendroecology of an old-growth, bottomland hemlock-white pine-northern harwood forest on the Allegheny Plateau. J. Torrey Bot. Soc. 126:234-244.

Orwig, D.A. and D.R. Foster. 2000. Stand, landscape, and ecosystem analyses of hemlock woolly adelgid outbreaks in southern New England: an overview. In Sustainable Management of Hemlock Ecosystems in Eastern North America. Symposium Proceedings USDA GTR-NE 267.

Yorks, T.E., J.C. Jenkins, D.J. Leopold, D.J. Raynal, and D.A. Orwig. 2000. Influences of eastern hemlock mortality on nutrient cycling. In Sustainable Management of Hemlock Ecosystems in Eastern North America. Symposium Proceedings USDA GTR-NE 267.

Orwig, D.A., C.V. Cogbill, D.R. Foster, and J.F. O'Keefe. 2001. Variations in old-growth structure and definitions: Development and dynamics of forests on Wachusett Mountain, Massachusetts. Ecological Applications 11:437-452.

Orwig, D.A. 2002a. Stand dynamics associated with chronic hemlock woolly adelgid infestations in southern New England. In Symposium on the Hemlock Woolly Adelgid in eastern North America proceedings. (eds. R.C. Reardon, B.P. Onken, and J. Lashomb) pp. 36-47. New Jersey Agricultural Experiment Station Publication, New Brunswick, NJ. In press.

Orwig, D.A. 2002b. Ecosystem to regional impacts of introduced pests and pathogens- historical context, questions, and issues. Journal of Biogeography 29: 1471-1474.

Cobb, R.C. and D.A. Orwig. 2002. Impact of hemlock woolly adelgid on decomposition: an overview. In Symposium on the Hemlock Woolly Adelgid in eastern North America proceedings. (eds. R.C. Reardon, B.P. Onken, and J. Lashomb) pp. 317-323. New Jersey Agricultural Experiment Station Publication, New Brunswick, NJ. In press.

Foster, D.R., S. Clayden, and D. Orwig, B. Hall, and S. Barry Musielewicz. 2002. Multiple-scale insights into the long-term history and stability of oak and chestnut-dominated hardwoods forests in Central New England. Journal of Biogeography 29: In press.

Motzkin, G., D.A. Orwig, and D.R. Foster. 2002. History and dynamics of a ridgetop pitch pine community: Mount Everett, Massachusetts. Harvard Forest Paper Number 25.

Orwig, D.A. and M.L. Kizlinski. 2002. Vegetation response following hemlock woolly adelgid infestation, hemlock decline, and hemlock salvage logging. In Symposium on the Hemlock Woolly Adelgid in eastern North America proceedings. (eds. R.C. Reardon, B.P. Onken, and J. Lashomb) pp. 106-118. New Jersey Agricultural Experiment Station Publication, New Brunswick, NJ. In press.

Orwig, D.A., D.R. Foster, and D.L. Mausel. 2002. Landscape Patterns of Hemlock Decline in New England Due to the Introduced Hemlock Woolly Adelgid. Journal of Biogeography 29: 1475-1487.

Kizlinski, M.L., D.A. Orwig, R.C. Cobb, and D.R. Foster. 2002. Direct and indirect ecosystem consequences of an invasive pest on forests dominated by eastern hemlock. J. of Biogeography 29: 1489-1503.

Motzkin, G., D.A. Orwig, and D.R. Foster. 2002. Vegetation and disturbance history of a rare dwarf pitch pine community on Mount Everett, Massachusetts. J. of Biogeography 29: 1455-1467.

Tingley, M.W., D.A. Orwig, G. Motzkin, D.R. Foster, and R. Field. 2002. Avian response to removal of a forest dominant: consequences of hemlock woolly adelgid infestations. J. of Biogeography 29: 1505-1515.

Foster, D. G. Motzkin, J. O’Keefe, E. Boose, D. Orwig, J. Fuller, and B. Hall. 2004. The environmental and human history of New England. In Forests in Time: The environmental consequences of 1,000 years of change in New England. Pp. 43-100. Yale University Press, New Haven, CT.

Stadler, B. T. Müller, D. Orwig, and R. Cobb. 2005. Hemlock woolly adelgid in New England forests: canopy impacts transforming ecosystem processes and landscapes. Ecosystems. In press.

Foster, D., D. Kittredge, B. Donahue, G. Motzkin, D. Orwig, A. Ellison, B. Hall, B. Colburn, and A. D’Amato. 2005. Wildlands and Woodlands: A vision for the forests of Massachusetts. Harvard Forest, Petersham, MA.

Stadler, B. T. Müller, D. Orwig, and R. Cobb. 2005. Hemlock woolly adelgid in New England forests: canopy impacts transforming ecosystem processes and landscapes. Ecosystems 8: 233-247.

Orwig, D.A. and D.B. Kittredge. 2005a. Silvicultural options for managing hemlock forests threatened by hemlock woolly adelgid. In: Reardon, R. and B. Onken (Eds). Proceedings of the 3^rd symposium on hemlock woolly adelgid in the eastern United States. USDA Forest Service, Newtown, PA. pp. 212-217.

Orwig, D.A. and D.B. Kittredge. 2005b. Silvicultural options for managing hemlock forests threatened by hemlock woolly adelgid. Umass extension fact sheet. Amherst, MA.

Jefts, S. and D. Orwig. 2005. The effects of HWA outbreaks on ecosystem level changes in southern New England. In: Reardon, R. and B. Onken (Eds). Proceedings of the 3^rd symposium on hemlock woolly adelgid in the eastern United States. USDA Forest Service, Newtown, PA. pp. 305-306.

Ellison, A.M., M.S. Bank, B.D. Clinton, E.A. Colburn, K. Elliott,^ C.R. Ford, D.R. Foster, B.D. Kloeppel, J.D. Knoepp, G.M. Lovett, J. Mohan, D.A. Orwig, N.L. Rodenhouse, W.V. Sobczak, K.A. Stinson, J.K. Stone, C.M. Swan, J. Thompson, B. Von Holle, and J.R. Webster. 2005. Loss of foundation species: consequences for the structure and dynamics of forested ecosystems. Frontiers in Ecology and the Environment. 3(9):479-486.

Foster. D.R. and D.A. Orwig. 2006. Pre-emptive and salvage harvesting of New England forests: when doing nothing is a viable alternative. Conservation Biology 20:959-970.

Stadler, B., T. Muller, and D. Orwig. 2006. The ecology of energy and nutrient fluxes in hemlock forests invaded by the hemlock woolly adelgid. Ecology 87:1792-1804.

Cobb. R.C, D.A. Orwig, and S. Currie. 2006. Decomposition of green foliage in eastern hemlock forests of southern New England impacted by hemlock woolly adelgid populations. Canadian Journal of Forest Research 36:1-11.

D’Amato, A.W., D.A. Orwig, and D.R. Foster. 2006. New estimates of Massachusetts old-growth forests: useful data for regional conservation and forest reserve planning. Northeastern Naturalist 13: 495-506.

Pederson. N., A.W. D’Amato, and D.A. Orwig. 2006. Central hardwood natural history from dendrochronology: maximum ages of rarely studied species. Proceedings of the 15^th Central Hardwood Forest Conference, Knoxville, TN. In press.

Preisser, E., A. Lodge, D. Orwig, and J. Elkinton. 2008. Range expansion and population dynamics of co-occurring invasive herbivores. Biological Invasions 10: 201-213.

Orwig, D.A., R.C. Cobb, A.W. D'Amato, M.L. Kizlinski, and D.R. Foster. 2008. Multi-year ecosystem response to hemlock woolly adelgid infestation in southern New England Forests. Canadian Journal of Forest Research 38: 834-843.

D'Amato, A.W., and D.A. Orwig. 2008. Stand and landscape-level disturbance dynamics in old-growth forest ecosystems in western Massachusetts. Ecologocial Monographs 78: 507-522.

D'Amato, A.W., D.A. Orwig, and D.R. Foster. 2008. The influence of successional processes and disturbance dynamics on the structure of old-growth and second-growth Tsuga canadensis forests. Ecological Applications 18: 1182-1199.

Cobb, R.C., and D.A. Orwig. 2008. Changes in decomposition dynamics in hemlock forests impacted by hemlock woolly adelgid: restoration and conservation of hemlock ecosystem function. In Proceedings: Fourth Symposium on hemlock woolly adelgid in the eastern United States ( eds. B. Onken and R. Reardon). pp 157-167. USDA Forest Service, Morgantown, WV.

Rollinson, C.R., and D.A. Orwig. 2008. Evaluating chemical treatments of hemlock woolly adelgid in southern New England. In Proceedings: Fourth Symposium on hemlock woolly adelgid in the eastern United States ( eds. B. Onken and R. Reardon). USDA Forest Service, Morgantown, WV. In press.

Smith, H.S., and D.A. Orwig. 2008. Influence of hemlock woolly adelgid and elongate hemlock scale on leaf-level physiological performance of eastern hemlock. In Proceedings: Fourth Symposium on hemlock woolly adelgid in the eastern United States ( eds. B. Onken and R. Reardon). USDA Forest Service, Morgantown, WV. In press.

Dukes, J.S., J. Pontius, D. Orwig, J.R. Garnas, V.L. Rodgers, N. Brazee, B. Cooke, K.A. Theoharides, E.E. Strange, R. Harrington, J. Ehrenfeld, J. Gurevitch, M. Lerdau, K. Stinson, R. Wick, and M. Ayres. 2009. Responses of insect pests, pathogens, and invasive plant species to climate change in the forests of Northeastern North America: What can we predict? Canadian Journal of Forest Research 39: 1-18.

D'Amato, D.A. Orwig, and D.R. Foster. 2009. Understory Vegetation in Old-Growth and Second-Growth Tsuga canadensis Forests in Western Massachusetts. Forest Ecology and Management 257: 1043-1052.